Couplet 4.
Male; antenna 14-segmented; F1 (and sometimes also F2) with a longitudinal carina, spine or emargination (Fig. 248-257); gaster with 7 free sternites
future couplets
Macropterous (Fig. 45) (p.52) (part) ...
Betyla
Gaster with 3 free tergites between... ...
Parabetyla
Gaster with fewer than 3 free tergi... ...
Betyla
Female; antenna 15-segmented; flagellar segments without carinae or emarginations (Fig. 239-247); gaster with 5 free sternites
future couplets
Tegula and mesoscutal-mesoscutellar... ...
Parabetyla
Tegula and mesoscutal-mesoscutellar... ...
Zealaptera
Gaster with only 1 completely free ... ...
Betyla